Great Demo, Dan. I listened to it LOUD.
It's a muff, but it's not really a muff. No diodes, no limiting resistors, different tone control. It's so much better than a muff.
I wonder if the difference in tone is down to component tolerances. Can you check the collector voltages on both?
EDIT: I remember now... the Raincoat is sensitive to HFE because there are no limiting resistors other than the SUSTAIN pot. Because there are no limiting resistors, the transistors are current-driven. That means that a higher HFE transistor can hit the next stage harder than a lower HFE transistor. The higher HFE results in a thicker tone and more sustain. 2N5133 and BC549C both have very broad HFE ranges and they mostly overlap. Unless you're hand-selecting transistors, you can't be sure what gain you have on any of them. You may even have a mix of high-gain and not-so-high-gain transistors on the same board.
If you want to try a way-out version of the Raincoat, take a look at this:
https://forum.pedalpcb.com/threads/quarantine-fuzz-viral-mutation.3229/
The Quarantine Fuzz is a Raincoat / Flugelhorn. I also built a Viral Mutation on a Flugelhorn board.