As I've mentioned before, I have very little understanding of the concept of impedance so take everything with a grain of salt.
Running some tests on LTspice, I've noticed that, when you plug a guitar straight to the input (assuming my pickup simulation is correct) there's a lot of treble roll off because of the low input impedance. However, when you feed the input from an ideal source (a buffer might act similarly), there's almost no treble cutting, which might be resulting in a lot of treble feeding the circuit down the road.
If that's the case, I'm not 100% sure Bean's suggestion will solve your problem.
Then again, I might be completely wrong here
Running some tests on LTspice, I've noticed that, when you plug a guitar straight to the input (assuming my pickup simulation is correct) there's a lot of treble roll off because of the low input impedance. However, when you feed the input from an ideal source (a buffer might act similarly), there's almost no treble cutting, which might be resulting in a lot of treble feeding the circuit down the road.
If that's the case, I'm not 100% sure Bean's suggestion will solve your problem.
Then again, I might be completely wrong here